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Genomic studies suggest modern wolves and dogs descend from a common ancestral wolf population that existed 20,000 years ago. Studies in 2017 and 2018 found that the Himalayan wolf is part of a lineage that is basal to other wolves and split from them 691,000–740,000 years ago. Other wolves appear to have originated in Beringia in an expansion that was driven by the huge ecological changes during the close of the Late Pleistocene. A study in 2016 indicates that a population bottleneck was followed by a rapid radiation from an ancestral population at a time during, or just after, the Last Glacial Maximum. This implies that the original morphologically diverse wolf populations were out-competed and replaced by more modern wolves.
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